Evolution, mutation (変異, hen'i) or transfiguration is the process by which one demon or Persona changes form into another, often the more powerful one that typically cannot be obtained via conventional means.
Shin Megami Tensei III: Nocturne[]
Certain demons can change into other demons by evolving. Some of these are unique in that they cannot be obtained through fusion in the Cathedral of Shadows, but all neutral demons in the list can appear in random encounters and be recruited normally. For a demon to evolve into a new form, it must learn all of its hidden skills from leveling up and Demi-fiend's level must be, at the minimum, four less than the resultant demon's. For example, High Pixie will not evolve into Queen Mab until Demi-fiend is at least level 52. If the result appears as a boss battle, that battle must be fought first (or, if the battle is avoidable due to the protagonist's Reason, pass the point of the story in which the fight normally occurs). When all requirements are met, an eligible demon will ask the protagonist's permission to evolve into its new form when it gains a level, even if the protagonist has declined it previously.
| Lvl | Race | Evolved from | > | Lvl | Race | Evolve into |
|---|---|---|---|---|---|---|
| 6 | Fairy | Pixie | 10 | Fairy | High Pixie# | |
| 15 | Fairy | High Pixie | 56 | Night | Queen Mab# | |
| 12 | Night | Lilim | 80 | Night | Lilith# | |
| 17 | Beast | Inugami | 22 | Avatar | Makami | |
| 18 | Snake | Nozuchi | 24 | Dragon | Gui Xian | |
| 22 | Beast | Nekomata | 28 | Holy | Senri | |
| 24 | Yoma | Koppa | 28 | Yoma | Karasu# | |
| 33 | Yoma | Karasu | 38 | Genma | Kurama | |
| 24 | Brute | Momunofu | 30 | Vile | Arahabaki | |
| 27 | Yoma | Dis | 33 | Wargod | Valkyrie | |
| 32 | Snake | Naga | 37 | Snake | Raja Naga# | |
| 38 | Snake | Mizuchi | 44 | Dragon | Long | |
| 42 | Yoma | Onkot | 46 | Genma | Hanuman | |
| 50 | Genma | Hanuman | 54 | Fury | Wu Kong | |
| 48 | Fairy | Setanta | 52 | Genma | Cu Chulainn | |
| 48 | Yoma | Jinn | 52 | Yoma | Efreet# | |
| 52 | Megami | Sati | 57 | Lady | Parvati | |
| 53 | Yoma | Purski | 58 | Wargod | Ganesha | |
| 58 | Beast | Sparna | 63 | Avian | Garuda | |
| 60 | Jirae | Gogmagog | 64 | Entity | Albion* | |
| 68 | Divine | Throne | 73 | Seraph | Uriel* | |
| 69 | Megami | Scathach | 74 | Lady | Skadi* | |
| 74 | Tyrant | Abaddon | 77 | Tyrant | Aciel* | |
| 86 | Tyrant | Beelzebub (human form) |
95 | Tyrant | Beelzebub (fly form)* |
- * Story boss
- # Neutral demon
The item "Divine Water," which can be purchased normally in shops, can be used to test if a demon can evolve, though this item is not cost-effective, as there are only 24 demons out of over 100 that can evolve. To ease the cost of checking whether or not his demons can evolve, the player can save before using it and reload after learning the result.
Shin Megami Tensei IV[]
Flynn's level is not a determining factor in whether or not a demon can evolve - a demon needs only to reach its own required level to trigger the event.
- Demons that form an evolution chain are highlighted in the same background color and can be sorted by the "Chain" column header in the table.
Shin Megami Tensei IV Apocalypse[]
The evolution chart remains mostly the same as the previous game, with a few minor changes due to the changes in availability of certain demons (for example, Inferno no longer evolves into Porewit, etc.). Demons gained through evolution will only have their base stats, rather than any higher stat levels the former form enjoyed. For example, a Girimehkala evolved from an Airavata with 106 Strength will only have 99 Strength.
- Demons that form an evolution chain are highlighted in the same background color and can be sorted by the "Chain" column header in the table.
| Lvl | Race | Evolved from | > | Race | Evolve into | Chain |
|---|---|---|---|---|---|---|
| 16 | Divine | Angel | Divine | Archangel | ||
| 22 | Divine | Archangel | Divine | Principality | ||
| 30 | Divine | Principality | Divine | Power | ||
| 37 | Divine | Power | Divine | Virtue | ||
| 46 | Divine | Virtue | Divine | Dominion | ||
| 56 | Divine | Dominion | Divine | Throne | ||
| 66 | Divine | Throne | Divine | Cherub | ||
| 78 | Divine | Cherub | Divine | Angel | ||
| 17 | Holy | Heqet | Lady | Dzelarhons | ||
| 21 | Beast | Kabuso | Nymph | Senri | ||
| 31 | Nymph | Senri | Beast | Nekomata | ||
| 20 | Yoma | Koppa Tengu | Yoma | Karasu Tengu | ||
| 23 | Spirit | Poltergeist | Spirit | Quicksilver | ||
| 30 | Spirit | Quicksilver | Wilder | Raijuu | ||
| 26 | Fairy | Jack Frost | Jaki | Wendigo | ||
| 27 | Vermin | Ubu | Jirae | Tsuchigumo | ||
| 28 | Ghost | Mou-Ryo | Spirit | Inferno | ||
| 34 | Femme | Yomotsu-Shikome | Brute | Yomotsu-Ikusa | ||
| 39 | Fairy | Kelpie | Yoma | Valkyrie | ||
| 40 | Night | Incubus | Night | Succubus | ||
| 42 | Undead | Corpse | Fallen | Nebiros | ||
| 50 | Avatar | Chimera | Beast | Orthrus | ||
| 64 | Beast | Orthrus | Beast | Cerberus | ||
| 52 | Drake | Mushussu | Drake | Tiamat | ||
| 60 | Drake | Tiamat | Drake | Ym | ||
| 57 | Holy | Airavata | Jaki | Girimehkala | ||
| 59 | Night | Wild Hunt | Tyrant | Abaddon | ||
| 62 | Vile | Tao Tie | Wilder | Taown | ||
| 70 | Femme | Dakini | Femme | Kali |
Last Bible II[]
Demons left at the beast training school can evolve into higher-leveled demons in their group.
Another Bible[]
Beasts evolve to a higher rank upon reaching level 3. Evolved beasts have 1 HP / PP more than beasts recruited at that rank, in addition to having the normal 5 stat points available upon leveling, making it preferable to evolve beasts to higher ranks rather than simply recruiting them. Rank 4 Beasts (Beast Fenrir, Fairy Oberon, and Tyrant Lucifer) can not be recruited, and are only available through evolution.
Devil Summoner: Soul Hackers[]
In the 3DS remake, the special demon Nemechi can evolve by consuming 300 D-Souls (once per day), which unlocks additional demons that can be obtained through it. The list of forms and purchasable demons is available here.
Persona 2[]
Many Personas have a small chance of mutating into another Persona of the minor Arcana of closest level after reaching Rank 8 and finishing a battle by performing a fusion spell. When a Persona shows signs of transformation, the player must take it to the Velvet Room and return it to the reserve stock.
There is a small chance for an evolution to result in a lvl. 2 Devil Poltergeist, which is considered a failed mutation. This can be prevented by reserving Poltergeist in either the protagonist's stock or in the Velvet Room before allowing a Persona to mutate, because the game does not allow the protagonist to possess multiple copies of any one Persona.
Innocent Sin[]
The following personas can mutate into higher level personas of the same arcana:
| Arcana | Lvl | Mutates from | Lvl | Mutates into |
|---|---|---|---|---|
| Any | Any | Any | 2 | Poltergeist |
| Magician | 41 | Abe no Seimei | 49 | Manannan |
| Priestess | 27 | Tensen Nyannyan | 34 | Sif |
| Hierophant | 15 | Xuanzang | 23 | Umayadono-Ouji |
| Hierophant | 53 | Mithra | 59 | Varuna |
| Hierophant | 59 | Varuna | 67 | Brahma * |
| Lovers | 2 | Pixie | 11 | Robin Goodfellow |
| Lovers | 11 | Robin Goodfellow | 18 | Jack Frost |
| Chariot | 43 | Susano-O | 50 | Ares |
| Hermit | 22 | Marshal Tianpeng | 28 | Gullinbursti |
| Hermit | 28 | Gullinbursti | 33 | Byakko |
| Hermit | 33 | Byakko | 40 | Genbu |
| Justice | 44 | Kundali Vidyaraja | 52 | Skanda |
| Hanged Man | 17 | Shax | 25 | Kabhanda |
| Death | 44 | Ankou | 47 | Charon * |
| Temperance | 54 | Phoenix | 59 | Suzaku * |
| Star | 15 | Kinnara | 18 | Iris |
| Star | 18 | Iris | 21 | Gandharva |
| Star | 60 | Valkyrie | 66 | Hanuman |
| Sun | 13 | Surya | 20 | Kinich Kakmo |
| Sun | 39 | Heimdall | 46 | Ildanach * |
| Judgement | 43 | Melchizedek | 48 | Armaiti |
| Judgement | 48 | Armaiti | 53 | Throne |
| World | 31 | Demeter | 38 | Njord * |
| World | 38 | Njord | 46 | Hunab Ku |
| World | 46 | Hunab Ku | 54 | Mucalinda |
| World | 54 | Mucalinda | 58 | Seiryu |
| World | 58 | Seiryu | 62 | Ouroboros |
Note:
* Indicates the Persona would normally require a Material Card to summon
Minor Arcana Mutations[]
Other Mutations[]
| Arcana | Mutates from | Lvl | Mutates into | Lvl | Remarks |
|---|---|---|---|---|---|
| Sun | Vulcanus | 1 | Vulcanus Prime | 33 | PSP only, if not obtained through dialogue choices; Complete Mt Iwato and Max Rank |
| Death | Rhadamanthus | 1 | Rhadamanthus Prime | 32 | |
| Lovers | Eros | 1 | Eros Prime | 32 | |
| Moon | Maia | 3 | Maia Prime | 31 | |
| Fortune | Hermes | 42 | Chronos | 57 | Complete Aquarius Temple and Max Rank |
Eternal Punishment[]
The following personas have a rather low chance to obtain via mutation.
| Lvl | Mutates from | Arcana | Lvl | Mutates into | Arcana | Remarks |
|---|---|---|---|---|---|---|
| 46 | Il-Dana | Sun | 72 | Lugh | Emperor | Requires rumor from Manannan |
| 64 | Scathach | Priestess | 74 | Pallas Athena | Justice | - |
| 68 | Seth | Tower | 74 | Mot | Death | - |
| 68 | Amurtart | Judgement | 83 | Michael | Judgement | 2nd Cycle |
| 68 | Wong Long | Strength | 88 | Shokuin | World | 2nd Cycle |
Minor Arcana Mutations[]
Other Mutations[]
| Arcana | Mutates from | Lvl | Mutates into | Lvl | Remarks |
|---|---|---|---|---|---|
| Moon | Maia | 3 | Maia Custom | 31 | Select correct dialogue in GOLD |
Persona 3[]
In Persona 3, the ultimate Personas of the protagonist's teammates evolve through story progression as a result of personal resolve. The protagonist's persona will evolve during the final battle with Nyx.
Persona 4[]
In Persona 4, Personas evolve into ultimate personas after an individual teammate's Social Link reaches max rank. The protagonist's persona will evolve during the final battle with Izanami.
Golden[]
In Persona 4 Golden, by talking to teammates on certain days after maxing out their social link, their ultimate persona will evolve into a third tier persona.
Persona 5[]
In Persona 5, Personas evolve into ultimate personas after an individual teammate's Confidant reaches max rank. The protagonist's persona will evolve during the final battle with Yaldabaoth.
Royal[]
In Persona 5 Royal, by talking to teammates on certain days after maxing out their Confidants, their ultimate persona will merge with their initial persona to evolve into a third tier persona.
In the Final Battle against Maruki, Azathoth evolves into Adam Kadmon.
Devil Children Black Book & Red Book[]
At certain points in the story Cool, Veil, and Nagahisa Kai's Sphinx will evolve into stronger forms. In Forest Land, the player will receive a phone call from Isis asking for them to visit her in Sand Land. Once there, she will give the player a Sphinx which can evolve by talking to a Trainer via the use of Emblems that can be found in certain dungeons. In Red Book, certain demons will evolve into a different one after being ranked up enough times.
Partners[]
| Evolved from | Evolve into | Requirement |
|---|---|---|
| Cerberus Gryphon |
Death Cerberus Death Gryphon |
Beat the Tournament in Marble Land. |
| Death Cerberus Death Gryphon |
Hell Cerberus Hell Gryphon |
Beat Nagahisa in Forest Land. (Black Book) Beat Nagahisa in Harajuku. (Red Book) |
| Hell Cerberus Hell Gryphon |
King Cerberus King Gryphon |
Rescue Lucifer in Central Land. |
Sphinx[]
| Evolved from | Evolve into | Requirement |
|---|---|---|
| Sphinx | Death Sphinx | Find Death Emblem in Forest Land. |
| Death Sphinx | Hell Sphinx | Find Hell Emblem in Central Land. |
| Hell Sphinx | Neo Sphinx | Find Neo Emblem in the Deep Hole. |
Rank up Evolutions[]
| Evolved from (Level) | Evolve into (Level) | Number of rank ups |
|---|---|---|
| Hippou (8) | Roc (17) | 5 |
| Deva (10) | Deva Ou (18) | |
| Bastet (13) | Neko Shogun (21) | |
| Feng Huang (19) | Suzaku (32) | |
| Komainu (20) | Yatsufusa (34) | |
| Shiisaa (28) | Barong (38) | 6 |
| Nekomata (5) | Bahan (14) | 5 |
| Shura (10) | Shura Ou (17) | |
| Tsukuyomi (44) | Scathach (50) | 8 |
| Van (49) | Van Ou (54) | 9 |
| Prince (23) | Virtue (32) | 6 |
| Kuriotes (39) | Merkabah (46) | 8 |
| Seraphim (45) | Archangel (52) | 9 |
| Ego (48) | Satan (54) | |
| Gnome (1) | Issun-boshi (9) | 5 |
| Pixie (3) | Sylph (16) | |
| Rasetsu (10) | Rasetsu Ou (17) | |
| Ghoul (24) | Mummy (32) | 6 |
| Robot (30) | Android (39) | |
| Counting Sheep (2) | Makura-gaeshi (13) | 5 |
| Makura-gaeshi (13) | Baku (23) | |
| Banshee (27) | Dullahan (40) | 6 |
| Mizuchi (8) | Otohime (14) | 5 |
| Basilisk (15) | Crom Cruach (27) | |
| Orochi (25) | Yamata no Orochi (34) | 6 |
| Naga (35) | Ananta (49) | 7 |
| Ratatoskr (3) | Kamaitachi (15) | 5 |
| Yukinko (7) | Yuki-onna (16) | |
| Kudagitsune (17) | Kyūbi no Kitsune (29) | |
| Frost Ace (22) | King Frost (31) | |
| Harpy (32) | Siren (41) | 7 |
Devil Children White Book[]
Clay and Rei will evolve as the story progresses.
DemiKids Light / Dark[]
Akira and Jin's demon partners Gale and Rand evolve as the story progresses. The choices made when the two are overflowing with power will determine the evolution path they follow. Rank Up fusion has several demons evolve into new demons by ranking up.
Devil Children Fire/Ice Book[]
Like in DemiKids: Light & Dark, Rand and Gale will once again evolve as the story progresses. This time however, the form they evolve into is based on whether their Attack or Magic stat is higher. The higher one leads down one path and it's possible to go down the other path if the opposite stat is higher by the next time the Demon partners can evolve. Rank Up fusion also returns and fulfills the same role it had in Dark/Light Version.
| Characters | |
| Playable | Masaki Kuzuha - Clay |
| Non-Playable | Takaharu Ougi - Rei - Shou Ougi - Paku - Haamiru - Setsuna Kai - Mirai Kaname - Cool - Veil - Takajo Zett - Nathanael - Raguel - Lucifer |
| Locations | |
| Earth | Harajuku |
| Expanse | Earth Expanse - Wind Expanse - Ice Expanse - Deep Hole - Center Expanse - Grass Field - Crystal Ring - Cosmos City - Altimia |
| Other | |
| Terms | Devil Riser - King Riser - Macca - Magnetite - Expanse - Devil Child - Angel Child - Devil Database - Devil Phone - Vinecom - Power Unit |
| Lists | Items - Demons - Skills |
| Light Version - Dark Version | |
| Characters | |
| Playable | Jin - Akira - Rand - Gale |
| Non-Playable | Lena - Amy Kashihara - Shin - Drail - Judy - Bul - Guy - Naomi - Lucifer - Darklord - Lord Light - Imperius - Imperius - Quazir - Seipher |
| Locations | |
| Rem | Rem Elementary School - Playground - Rem Mall - Rem Park - Rem Blvd. |
| Future Rem | Rem Mall (Future) - Rem Blvd. (Future) - Rem Park (Future) |
| Redwood | Rem Mall (Future) - Sinra Forest - Redwood Town |
| DemWind | Hueyville - Gateway - Wind Valley - Sword Cave - Kibra Woods - Wind Shrine - Wind Gate - Lost Forest - Galin Plains |
| DemCenter | Center Town - Heldun Flats - Dark Palace - Orgel Room - Deep Hole |
| DemIce | Frostville - Polar Wastes - Ace Base - Ice Cave - Ice Temple - Leader Home - Icy Wastes - Luna Prairie |
| DemSub | Peril Forest - Underworld - Annwn Town |
| DemFire | Helkane Peak - Volkanville - Mt. Giles - Myorl Town - Dark Path - Dark Temple |
| DemSand | Valk Desert - Nobleville - Dune Desert - Howling - Light Path - Light Temple |
| Valhalla | Time's Edge - Time Tower - Tower Area - Fanda Plains - Hermit - Duvegs Trail - Hydraville - Hydra Mine - Limbo - Inferno Pit - Foggy Marsh - Lair Area - Rebel's Lair - Jude Plains - Mount Falos - Elysium - Avalon - Rainy Woods - Tyrnanog |
| Archives | |
| Terms | Demiloc - Kingloc - Vinecom - DemiKid - Vinecenter - Battlenet |
| Lists | Demons - Skills - Powers - Items |
| Other Media | |
| Productions | Animation (Episodes) |
| Publications | Manga |
| Characters | |
| Playable | Jin - Akira - Rand - Gale |
| Non-Playable | Lena - Amy Kashihara - Paku - Haamiru - Mephisto - Isaac - Michael - Lucifer - Faruha - Kerumido - Seku - Rejiinu - Ganku - Kuun - Muuru - Flame Apostle - Ice Apostle - Shugoshin - Secundus - Zosma |
| Locations | |
| Harajuku | Main Street - Park - Mansion - Underground Shopping Center - School - Akuma no Hokora - Angel's Tower |
| Makai | Inelus Village - Acheron River - Central Land - Dark Palace - Deep Hole - Black Forest - Peril Forest - Underworld - Annwn Town - Heimdall Plains - Babylon Labyrinth |
| Tenkai | Midkingdom - Highest Peak - Light Palace - Fire Machine Mountain - Eden - Huge Floodgate - Water Angels Village - Lotus Marsh - Dios Forest - Earth Angels Village - Extensive Earthly Forest - Tower of Babel - Seele Plains - Shugyōja Road - Wind Angels Village - Fire Angels Village |
| Niflheim | Suneefiyoru - Dark North - Hvergelmir - Gimle Snowfield - Norðri Shrine - Niðafjöll - Vestri Shrine - Ice Apostle Castle - Ice Apostles Castle Town |
| Muspelheim | Soorufiyoru - Austri Shrine - Mímisbrunnr - Fairy Village - Rogafieru - Suðri Shrine - Flame Apostle Castle - Flame Apostles Castle Town |
| Ginnungagap | Auðumla Town - Huge Bridge - Bridge Cave - Shugoshin Cave |
| Other | |
| Lists | Demons - Skills |