Reverse Persona

Reverse Personas are the negative form of the original Personas. They are usually used by the boss characters in the battle. The concept exists exclusively in the Persona 2 duology but is implied in Persona 4.

Persona 2: Innocent Sin
Reverse Personas can be divided into to 2 category, one is used by the members of the Masked Circle, they are:
 * Reverse Rhadamanthus by Hidehiko Uesugi
 * Reverse Shax by Yasuo Inoe
 * Reverse Eros by Ginji Sasaki
 * Reverse Parica by Ixquic
 * Reverse Vulcanus by Tatsuya Sudou

Presumably Youichi Makimura possessed a Reverse Persona as well but the actual Persona he uses is never shown on-screen. Junko Kurosu, being one of the Masked Circle executives, whose ability to summon a Persona is unknown.

The other group of the Reverse Personas are used by the Shadow Selves of the party members:
 * Reverse Maia by Shadow Maya
 * Reverse Durga by Shadow Yukino
 * Reverse Apollo by Shadow Tatsuya
 * Reverse Venus by Shadow Ginko
 * Reverse Hades by Shadow Eikichi

It can be theorized that Jun Kurosu's Persona when assuming his Joker identity, Joker-Nyarlathotep, is his Reverse Persona. This is supported by fact that all Reverse Personas and Shadow Selves are creations of the shadow of humanity, in other word, Nyarlathotep itself.