List of Digital Devil Saga: Avatar Tuner Bosses

Svadhisthana (Vanguards Base)

 * Hayagriva (Avatar form of Harley)

Manipura (Maribel Base)

 * Camazotz (Avatar form of Bat)

Anahata (Solids Citadel)

 * Usas (Avatar form of Jinana)

Manipura (Maribel Base), Second Time

 * Camazotz (Avatar form of Bat)
 * Atavaka

Coordinate 136

 * Rahu (Avatar form of Mick the Slug)

Deserted Ship

 * Camaztotz (Heaven or Hell) (Avatar form of Bat)

Svadhisthana Waterways

 * Cerberus (Avatar form of Lupa)

Ajna (Brutes Base)

 * Catoblepas
 * Isis
 * Nidhoggr
 * Ravana (Avatar form of Varin Omega)

Karma Temple

 * Vasuki
 * Jatayu
 * Garuda
 * Ananta
 * Harihara (Onmyo form of Angel)

Svadhisthana (Vanguards Base)

 * Human Beelzebub
 * Feng Huang of the Four Guardian Beasts

Manipura (Maribel Base)

 * Baihu of the Four Guardian Beasts

Manipura Waterways

 * Fly Beelzebub

Anahata (Solids Citadel)

 * Gui Xian of the Four Guardian Beasts
 * Demi-Fiend

Anahata Waterways

 * Orochi

Coordinate 136

 * King Frost

Ajna (Brutes Base)

 * Long of the Four Guardian Beasts
 * Metatron

Karma Temple

 * Huang Long